I'd be interested in hearing more about how you define "culture" among species. In your article, you seem to define culture as how apes invent and use tools. Are there other ways of defining culture? I suppose how they organize themselves, which family member does child care, and how they protect themselves is considered anthopology rather than culture. Also, orangutan tool use has proven to reveal some of the most unique functions in the entire animal kingdom, including autoerotic tool-use, leaf napkin, branch swatter, seed extraction tool-use, sun cover (building a cover for a nest on bright sunny days) and branch scoop (drinking water from a deep tree hole using a leafy branch) (van Schaik et al., 2003).
Hey lil, When primatologists (and other animal behaviourists) use the term culture it is usually to describe behaviour and behavioural variation that is caused by social learning (as opposed to genetic or ecologically determined behaviour). And also, I am completely intrigued with what future studies will discover regarding orangutan and gorilla cultural variation. Already with little research there have been some really interesting discoveries.